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Atlas tissue group Human Homo sapiens

heart muscle

The Human Protein Atlas classes 429 genes as elevated in heart muscle (32 tissue enriched, 126 group enriched, 271 tissue enhanced), in Human Protein Atlas version 25.1 (release 2026-05-25); entry tag 25, read 2026-09-09. The values are the atlas's own nTPM, which is not TPM; nothing on this page is compared with a GTEx value.

The atlas's own pages

The elevated genes came from the atlas's search field for the identical expression the link below carries, so the page it opens lists the rows this one holds.

The same search at the atlas

01The elevated genes

Genes the atlas classes as elevated in heart muscle

What this tells you

The genes are the atlas's own specificity classification for this tissue group, read from its search field on 2026-09-09 and built into this site. The atlas release is version 25.1 (release 2026-05-25); entry tag 25: the entry tag is read from the one machine-readable statement the atlas makes of its version, the entry tag of its per-gene record, in the same build, and the version number and release date beside it are the atlas's release history page's [R30], quoted below. The consensus card is the atlas's published consensus table, cut to its largest values for this tissue.

The categories, as the atlas defines them [R29]. Enriched: nTPM in a particular tissue/region/cell type at least four times any other tissue/region/cell type [R29]. Group enriched: nTPM in a group (of 2-5 tissues, brain regions, single cell types or cell lines, or 2-10 immune cell types) at least four times any other tissue/region/cell line/immune cell type/cell type [R29]. Enhanced: nTPM in a one or several tissues, brain regions, cell lines, immune cell types or single cell types that has at least four times the mean of all tissue/region/cell types [R29]. The score beside an enriched or group-enriched gene: TS/CS-score is calculated as the fold change from the tissue/cell line with highest RNA to the tissue/cell line with second highest RNA. [R29] The atlas publishes none for a tissue-enhanced gene, and the table shows none.

The unit is the atlas's own: all TPM values of all samples within each data source (HPA + GTEx human tissues, HPA immune cell types, HPA cell lines) were normalized separately using Trimmed mean of M values (TMM) to allow for between-sample comparisons. The resulting normalized transcript expression values, denoted nTPM, were calculated for each gene in every sample. nTPM values below 0.1 are not visualized on the Atlas sections. [R29] nTPM is not TPM, and no value here is put beside a GTEx value from this site's GTEx pages. The consensus value is a maximum, never an average: The consensus nTPM value for each gene and tissue type represents the maximum nTPM value based on HPA and GTEx. For tissues with multiple sub-tissues (brain regions, immune cells, lymphoid tissues and intestine) the maximum of all sub-tissues is used for the tissue type and the total number of tissue types in the human tissue consensus set is 37. [R29]

The release: Protein Atlas version 25.1. Release date: 2026.05.25. Ensembl version: 109. [R30] Over the whole atlas, its tissue resource says all putative 20162 protein coding genes have been classified with regard to abundance and distribution of transcribed mRNA molecules, including 11035 proteins showing a significantly elevated level of expression in a particular tissue or a group of related tissues and 8813 proteins detected in all organs and tissues [R31], and its specificity counts on that page are 3132 tissue enriched, 1547 group enriched, 6356 tissue enhanced, 8096 of low tissue specificity and 1031 not detected [R31]. The atlas is licensed under the Creative Commons Attribution 4.0 International License for all copyrightable parts of our database [R28] and asks a website to cite the source in a manner that is clear, accurate and easily discoverable and link to the source [R28], which the provenance line under each card does, naming its primary publication [R07] and the versioned address the data came from.

An elevated gene here is one the atlas classes as elevated in this group by its own thresholds over its own consensus values, of which the GTEx column is GTEx v8 folded into nTPM; a value on this page and a TPM on this site's GTEx pages are two measurements of two sample sets in two units.

  1. [R07] Uhlén M, Fagerberg L, Hallström BM, Lindskog C, Oksvold P, Mardinoglu A, et al. (2015). Tissue-based map of the human proteome. Science 347:1260419. PMID 25613900, doi 10.1126/science.1260419.
  2. [R28] The Human Protein Atlas, proteinatlas.org. Licence & Citation. https://www.proteinatlas.org/about/licence, read 2026-09-09.
  3. [R29] The Human Protein Atlas, proteinatlas.org. The human proteome, Methods summary, Transcriptomics. https://www.proteinatlas.org/humanproteome/tissue/method/transcriptomics, read 2026-09-09.
  4. [R30] The Human Protein Atlas, proteinatlas.org. Release history. https://www.proteinatlas.org/about/releases, read 2026-09-09.
  5. [R31] The Human Protein Atlas, proteinatlas.org. Tissue resource, Tissue-based map of the human proteome. https://www.proteinatlas.org/humanproteome/tissue, read 2026-09-09.

The atlas classes 126 genes as group enriched in heart muscle; showing 1 to 100 in pages of 100, in the atlas's own order. The nTPM is the atlas's own unit.

Genes the Human Protein Atlas classes as group enriched in heart muscle, page 1 of 2
GeneACTN2EnsemblENSG00000077522TS-score35nTPM695.9Elevated inheart muscle 695.9; skeletal muscle 1,961.6; tongue 1,195.2
GeneADPRHL1EnsemblENSG00000153531TS-score32nTPM274.1Elevated inheart muscle 274.1; skeletal muscle 246.1; tongue 213.8
GeneANKRD2EnsemblENSG00000165887TS-score44nTPM495.5Elevated inheart muscle 495.5; skeletal muscle 1,141.9; tongue 769.1
GeneANO5EnsemblENSG00000171714TS-score8nTPM22.7Elevated inheart muscle 22.7; parathyroid gland 16.2; skeletal muscle 61.5; tongue 46.5
GeneART3EnsemblENSG00000156219TS-score10nTPM63.7Elevated inheart muscle 63.7; skeletal muscle 190.7; testis 86.3; tongue 70.8
GeneASB11EnsemblENSG00000165192TS-score14nTPM31.3Elevated inheart muscle 31.3; skeletal muscle 107.3; tongue 57.4
GeneASB18EnsemblENSG00000182177TS-score6nTPM6.6Elevated inheart muscle 6.6; skeletal muscle 11
GeneASIPEnsemblENSG00000101440TS-score5nTPM23.2Elevated inepididymis 11.5; heart muscle 23.2; ovary 34.5
GeneATP1A3EnsemblENSG00000105409TS-score8nTPM147.7Elevated inbrain 279; heart muscle 147.7; retina 271.7
GeneATP2A2EnsemblENSG00000174437TS-score12nTPM941.9Elevated inheart muscle 941.9; skeletal muscle 2,675.7; tongue 2,586.2
GeneC11orf21EnsemblENSG00000110665TS-score6nTPM19.4Elevated inbone marrow 47.6; heart muscle 19.4; lymphoid tissue 15
GeneC19orf47EnsemblENSG00000160392TS-score6nTPM164.2Elevated inheart muscle 164.2; skeletal muscle 335.4; tongue 128.4
GeneC1orf105EnsemblENSG00000180999TS-score4nTPM16.8Elevated inheart muscle 16.8; testis 24.6
GeneCAMK2BEnsemblENSG00000058404TS-score6nTPM62.4Elevated inbrain 104.5; heart muscle 62.4; skeletal muscle 112.2; tongue 57.4
GeneCDH19EnsemblENSG00000071991TS-score4nTPM35.4Elevated inblood vessel 14.1; heart muscle 35.4; intestine 18.6
GeneCDIN1EnsemblENSG00000186073TS-score5nTPM73.6Elevated inheart muscle 73.6; tongue 32.2
GeneCDK18EnsemblENSG00000117266TS-score7nTPM116.7Elevated inbrain 367.7; heart muscle 116.7
GeneCFL2EnsemblENSG00000165410TS-score5nTPM348.8Elevated inheart muscle 348.8; skeletal muscle 674.8; tongue 640.8
GeneCHRNEEnsemblENSG00000108556TS-score9nTPM57.6Elevated inheart muscle 57.6; pituitary gland 27.7
GeneCKMT2EnsemblENSG00000131730TS-score28nTPM1,157.4Elevated inheart muscle 1,157.4; skeletal muscle 1,341.7; tongue 2,166.2
GeneCOQ10AEnsemblENSG00000135469TS-score7nTPM157.6Elevated inheart muscle 157.6; skeletal muscle 263.1; tongue 361.4
GeneCOX6A2EnsemblENSG00000156885TS-score41nTPM4,887.8Elevated inheart muscle 4,887.8; skeletal muscle 6,326; tongue 1,952.3
GeneCOX7A1EnsemblENSG00000161281TS-score12nTPM4,516.8Elevated inheart muscle 4,516.8; skeletal muscle 6,519.5; tongue 4,622.3
GeneCPLX3EnsemblENSG00000213578TS-score5nTPM44.8Elevated inbrain 69.2; heart muscle 44.8; lymphoid tissue 117.6; prostate 127.2; retina 91.5
GeneCPNE5EnsemblENSG00000124772TS-score4nTPM90Elevated inbrain 131.7; heart muscle 90
GeneCPT1BEnsemblENSG00000205560TS-score4nTPM226.2Elevated inheart muscle 226.2; skeletal muscle 136.9; testis 75.9; tongue 212.9
GeneCYP2J2EnsemblENSG00000134716TS-score4nTPM83.9Elevated inheart muscle 83.9; intestine 78.5; liver 136.7
GeneDHRS7CEnsemblENSG00000184544TS-score64nTPM102.3Elevated inheart muscle 102.3; skeletal muscle 288.6; tongue 342.6
GeneDTNAEnsemblENSG00000134769TS-score4nTPM140.1Elevated inbrain 204.2; choroid plexus 72; heart muscle 140.1; skeletal muscle 152.8; tongue 197.5
GeneENSG00000278817EnsemblENSG00000278817TS-score5nTPM51.8Elevated inchoroid plexus 75; heart muscle 51.8; skeletal muscle 107; tongue 125.6
GeneFABP3EnsemblENSG00000121769TS-score12nTPM1,397.8Elevated inheart muscle 1,397.8; skeletal muscle 525.2; tongue 1,347.3
GeneFBXO40EnsemblENSG00000163833TS-score96nTPM84.7Elevated inheart muscle 84.7; skeletal muscle 152; tongue 93.8
GeneFHL2EnsemblENSG00000115641TS-score8nTPM1,337.6Elevated inheart muscle 1,337.6; ovary 513.6
GeneFITM1EnsemblENSG00000139914TS-score7nTPM163.8Elevated inheart muscle 163.8; skeletal muscle 218.2
GeneFSD2EnsemblENSG00000186628TS-score35nTPM49Elevated inheart muscle 49; skeletal muscle 144.6; tongue 41.4
GeneGABRA4EnsemblENSG00000109158TS-score7nTPM2.4Elevated inbrain 9.3; heart muscle 2.4
GeneGOT2EnsemblENSG00000125166TS-score5nTPM297.9Elevated inchoroid plexus 205.8; heart muscle 297.9; liver 300.5; skeletal muscle 654.1; tongue 678.6
GeneGRIK2EnsemblENSG00000164418TS-score5nTPM4Elevated inbrain 10; heart muscle 4
GeneGRIP2EnsemblENSG00000144596TS-score5nTPM12.4Elevated inheart muscle 12.4; skeletal muscle 33.5; tongue 11.6
GeneGSG1LEnsemblENSG00000169181TS-score9nTPM21.6Elevated inblood vessel 18.1; brain 19.8; heart muscle 21.6
GeneGYS1EnsemblENSG00000104812TS-score5nTPM112.2Elevated inheart muscle 112.2; skeletal muscle 299.1; tongue 188.4
GeneHCN4EnsemblENSG00000138622TS-score6nTPM8.2Elevated inheart muscle 8.2; testis 7
GeneHHATLEnsemblENSG00000010282TS-score8nTPM487.7Elevated inbrain 233.4; heart muscle 487.7; skeletal muscle 264.6; tongue 144
GeneHRCEnsemblENSG00000130528TS-score26nTPM692.1Elevated inheart muscle 692.1; skeletal muscle 424.2; tongue 228.2
GeneHSPB3EnsemblENSG00000169271TS-score19nTPM463.9Elevated inheart muscle 463.9; skeletal muscle 232.7; tongue 204.1
GeneHTR4EnsemblENSG00000164270TS-score5nTPM4.3Elevated inbrain 2.1; heart muscle 4.3; intestine 8.5
GeneIDH2EnsemblENSG00000182054TS-score4nTPM462.3Elevated inheart muscle 462.3; skeletal muscle 1,220.1; tongue 1,504.2
GeneISLR2EnsemblENSG00000167178TS-score4nTPM4.4Elevated inbrain 15.5; heart muscle 4.4; testis 12.3
GeneKCNA5EnsemblENSG00000130037TS-score6nTPM36.5Elevated inblood vessel 42.2; choroid plexus 57.3; heart muscle 36.5
GeneKCNE1EnsemblENSG00000180509TS-score5nTPM10.7Elevated inchoroid plexus 14.9; fallopian tube 14.4; heart muscle 10.7; kidney 3.9
GeneKCNJ4EnsemblENSG00000168135TS-score18nTPM31.7Elevated inbrain 82.7; heart muscle 31.7
GeneLDB3EnsemblENSG00000122367TS-score24nTPM1,170.6Elevated inheart muscle 1,170.6; skeletal muscle 2,717.5; tongue 1,686.1
GeneLMAN1LEnsemblENSG00000140506TS-score6nTPM31.5Elevated inbrain 24.1; heart muscle 31.5; lymphoid tissue 24.2; prostate 57.2; salivary gland 19.5
GeneLMOD2EnsemblENSG00000170807TS-score39nTPM728.6Elevated inheart muscle 728.6; skeletal muscle 874.4; tongue 741.3
GeneLRRC14BEnsemblENSG00000185028TS-score4nTPM39.2Elevated inheart muscle 39.2; skeletal muscle 106.6
GeneLRRC2EnsemblENSG00000163827TS-score5nTPM42.9Elevated inheart muscle 42.9; skeletal muscle 154.3; tongue 88.2
GeneMBEnsemblENSG00000198125TS-score38nTPM14,050.6Elevated inheart muscle 14,050.6; skeletal muscle 38,156.8; tongue 27,139.2
GeneMLIPEnsemblENSG00000146147TS-score10nTPM186.2Elevated inheart muscle 186.2; liver 119.5; skeletal muscle 205.1; tongue 139.5
GeneMPP3EnsemblENSG00000161647TS-score6nTPM14.5Elevated inbrain 28.2; heart muscle 14.5
GeneMTLNEnsemblENSG00000175701TS-score5nTPM232.8Elevated inheart muscle 232.8; skeletal muscle 736.2; tongue 262.1
GeneMTUS2EnsemblENSG00000132938TS-score4nTPM47Elevated inblood vessel 12.9; heart muscle 47
GeneMYH7EnsemblENSG00000092054TS-score71nTPM7,145.3Elevated inheart muscle 7,145.3; skeletal muscle 13,633.1; tongue 5,024.6
GeneMYH7BEnsemblENSG00000078814TS-score9nTPM338.7Elevated inheart muscle 338.7; skeletal muscle 147.2
GeneMYL12AEnsemblENSG00000101608TS-score5nTPM3,965.3Elevated inheart muscle 3,965.3; skeletal muscle 3,053.6; tongue 1,943.8
GeneMYL2EnsemblENSG00000111245TS-score73nTPM27,634.5Elevated inheart muscle 27,634.5; skeletal muscle 27,153.8; tongue 18,492.7
GeneMYLK3EnsemblENSG00000140795TS-score13nTPM96.8Elevated inheart muscle 96.8; skeletal muscle 43.4; tongue 67.7
GeneMYOM1EnsemblENSG00000101605TS-score18nTPM375.8Elevated inheart muscle 375.8; skeletal muscle 762.9; tongue 399.9
GeneMYOM2EnsemblENSG00000036448TS-score6nTPM324.6Elevated inheart muscle 324.6; skeletal muscle 372
GeneMYOM3EnsemblENSG00000142661TS-score9nTPM209.7Elevated inheart muscle 209.7; skeletal muscle 308.7; tongue 238.9
GeneNDUFB10EnsemblENSG00000140990TS-score5nTPM755.8Elevated inheart muscle 755.8; skeletal muscle 1,217.5; tongue 945.6
GeneNDUFC1EnsemblENSG00000109390TS-score6nTPM78.4Elevated inheart muscle 78.4; skeletal muscle 251.3; tongue 148.5
GeneNIPSNAP2EnsemblENSG00000146729TS-score7nTPM249.4Elevated inheart muscle 249.4; skeletal muscle 604.5; tongue 732.5
GeneNKX2-6EnsemblENSG00000180053TS-score13nTPM1Elevated inheart muscle 1; salivary gland 1.5
GeneNMRK2EnsemblENSG00000077009TS-score88nTPM1,378.3Elevated inheart muscle 1,378.3; skeletal muscle 590.6; tongue 669.1
GeneOTOGLEnsemblENSG00000165899TS-score4nTPM4.1Elevated inheart muscle 4.1; intestine 1.5; kidney 4.8
GenePACSIN3EnsemblENSG00000165912TS-score4nTPM155.2Elevated inheart muscle 155.2; skeletal muscle 450.8; tongue 260.7
GenePDE4DIPEnsemblENSG00000178104TS-score8nTPM686.9Elevated inheart muscle 686.9; skeletal muscle 1,934.8; tongue 1,153.3
GenePDLIM5EnsemblENSG00000163110TS-score7nTPM589.2Elevated inheart muscle 589.2; skeletal muscle 1,089.9; tongue 538
GenePERM1EnsemblENSG00000187642TS-score56nTPM64.2Elevated inheart muscle 64.2; skeletal muscle 168.4; tongue 126.9
GenePGAM2EnsemblENSG00000164708TS-score13nTPM1,112.4Elevated inheart muscle 1,112.4; skeletal muscle 4,078.1; tongue 1,636.7
GenePLPP7EnsemblENSG00000160539TS-score7nTPM87.4Elevated inheart muscle 87.4; skeletal muscle 196.8; tongue 69.5
GenePOPDC3EnsemblENSG00000132429TS-score6nTPM19.4Elevated inheart muscle 19.4; skeletal muscle 76.4; tongue 27.6
GenePPP1R3AEnsemblENSG00000154415TS-score63nTPM38.2Elevated inheart muscle 38.2; skeletal muscle 143.1; tongue 97.2
GenePPP2R3AEnsemblENSG00000073711TS-score5nTPM47.9Elevated inheart muscle 47.9; skeletal muscle 168.9; tongue 110
GenePPP2R3BEnsemblENSG00000167393TS-score5nTPM59.2Elevated inheart muscle 59.2; skeletal muscle 87.3
GeneRBM24EnsemblENSG00000112183TS-score6nTPM132.9Elevated inheart muscle 132.9; skeletal muscle 293.2; tongue 98.7
GeneRD3LEnsemblENSG00000227729TS-score31nTPM21.8Elevated inheart muscle 21.8; retina 12.4
GeneRRADEnsemblENSG00000166592TS-score5nTPM316.2Elevated inheart muscle 316.2; skeletal muscle 776.8
GeneS100A1EnsemblENSG00000160678TS-score5nTPM2,882.1Elevated inbrain 1,266.4; heart muscle 2,882.1; skeletal muscle 2,868.7; tongue 4,577.6
GeneSCHIP1EnsemblENSG00000151967TS-score8nTPM74.5Elevated inbrain 76.6; heart muscle 74.5; skeletal muscle 92.5; tongue 86.3
GeneSFRP5EnsemblENSG00000120057TS-score4nTPM71.7Elevated inchoroid plexus 215.9; heart muscle 71.7; pancreas 135.1
GeneSGCAEnsemblENSG00000108823TS-score5nTPM118.7Elevated inblood vessel 168.4; heart muscle 118.7; skeletal muscle 435.8; tongue 202.7
GeneSGCGEnsemblENSG00000102683TS-score10nTPM111.2Elevated inheart muscle 111.2; skeletal muscle 148; tongue 134.5
GeneSHDEnsemblENSG00000105251TS-score7nTPM21.3Elevated inbrain 12.7; heart muscle 21.3
GeneSLC25A11EnsemblENSG00000108528TS-score4nTPM218.8Elevated inheart muscle 218.8; skeletal muscle 294.7; tongue 419.8
GeneSLC25A34EnsemblENSG00000162461TS-score4nTPM13.4Elevated inbrain 15.3; heart muscle 13.4; intestine 13; skeletal muscle 36.5; tongue 43.5
GeneSLC25A4EnsemblENSG00000151729TS-score5nTPM240.1Elevated inheart muscle 240.1; skeletal muscle 191.3; tongue 323.8
GeneSMCO1EnsemblENSG00000214097TS-score41nTPM39.4Elevated inheart muscle 39.4; skeletal muscle 139.7; tongue 99.9
GeneSMIM10L2BEnsemblENSG00000196972TS-score4nTPM10.2Elevated inbrain 38.2; heart muscle 10.2
GeneSMPXEnsemblENSG00000091482TS-score38nTPM709.4Elevated inheart muscle 709.4; skeletal muscle 1,108.9; tongue 1,025.1

TS-score is the atlas's tissue specificity score, published for enriched and group-enriched genes and for no tissue-enhanced gene; none published is the atlas's absence, not a zero. The nTPM column is the value the atlas labels heart muscle; a group-enriched gene lists every group the atlas names for it, with the atlas's own labels.

Pages of 100 are this site's own cut of the atlas's answer, which came whole; the categories are the three elevated categories the atlas defines, and its other two, low tissue specificity and not detected, are not tissue lists and are not shown.

  • Human Protein Atlas, the tissue specificity field for heart muscle · Human Protein Atlas version 25.1 (release 2026-05-25); entry tag 25 · read · the same search at the atlasHuman Protein Atlas version 25.1 (release 2026-05-25); entry tag 25, proteinatlas.org; data available from v25.proteinatlas.org/search/tissue_category_rna:heart+muscle;tissue+enriched,group+enriched,tissue+enhanced (Uhlén M et al. Science 2015). CC BY 4.0.

02The consensus values

The atlas's consensus nTPM for heart muscle

The atlas's consensus table carries 20,162 rows for heart muscle, 16,738 of them above zero; showing the 250 largest values, which is this site's own cut of the table, in nTPM, the atlas's own unit.

The consensus nTPM value for each gene and tissue type represents the maximum nTPM value based on HPA and GTEx. For tissues with multiple sub-tissues (brain regions, immune cells, lymphoid tissues and intestine) the maximum of all sub-tissues is used for the tissue type and the total number of tissue types in the human tissue consensus set is 37.

A consensus nTPM of 0.0 in this table means below 0.1 nTPM, which is the smallest value the atlas publishes here. It does not mean the gene was not detected.

The 250 largest consensus nTPM values the Human Protein Atlas publishes for heart muscle
Rank1GeneMT-ATP8EnsemblENSG00000228253nTPM355,486.2
Rank2GeneMT-CO1EnsemblENSG00000198804nTPM265,338.6
Rank3GeneMT-ATP6EnsemblENSG00000198899nTPM240,225.6
Rank4GeneMT-CO3EnsemblENSG00000198938nTPM238,539.3
Rank5GeneMT-CO2EnsemblENSG00000198712nTPM214,510.8
Rank6GeneMT-ND4EnsemblENSG00000198886nTPM197,808.9
Rank7GeneMT-ND3EnsemblENSG00000198840nTPM186,829.7
Rank8GeneMT-ND4LEnsemblENSG00000212907nTPM149,995.1
Rank9GeneMT-CYBEnsemblENSG00000198727nTPM131,222.4
Rank10GeneMT-ND2EnsemblENSG00000198763nTPM130,852.9
Rank11GeneMT-ND1EnsemblENSG00000198888nTPM129,323.4
Rank12GeneNPPAEnsemblENSG00000175206nTPM57,628
Rank13GeneMT-ND6EnsemblENSG00000198695nTPM33,513
Rank14GeneMYL2EnsemblENSG00000111245nTPM27,634.5
Rank15GeneMT-ND5EnsemblENSG00000198786nTPM26,164.9
Rank16GeneMYL7EnsemblENSG00000106631nTPM22,266.8
Rank17GeneMBEnsemblENSG00000198125nTPM14,050.6
Rank18GeneACTC1EnsemblENSG00000159251nTPM10,840.1
Rank19GeneCRYABEnsemblENSG00000109846nTPM10,537.6
Rank20GeneTPM1EnsemblENSG00000140416nTPM10,511.2
Rank21GeneTNNI3EnsemblENSG00000129991nTPM10,359.2
Rank22GeneMYL3EnsemblENSG00000160808nTPM8,764.4
Rank23GeneTNNC1EnsemblENSG00000114854nTPM8,750.1
Rank24GeneTNNT2EnsemblENSG00000118194nTPM8,731.9
Rank25GeneMYH7EnsemblENSG00000092054nTPM7,145.3
Rank26GeneTCAPEnsemblENSG00000173991nTPM6,421.7
Rank27GeneACTA1EnsemblENSG00000143632nTPM6,223.8
Rank28GeneANKRD1EnsemblENSG00000148677nTPM6,125.5
Rank29GeneMYL4EnsemblENSG00000198336nTPM6,049.7
Rank30GeneDESEnsemblENSG00000175084nTPM5,337.8
Rank31GeneGAPDHEnsemblENSG00000111640nTPM5,080.5
Rank32GeneCOX6A2EnsemblENSG00000156885nTPM4,887.8
Rank33GeneMYH6EnsemblENSG00000197616nTPM4,840.4
Rank34GeneNPPBEnsemblENSG00000120937nTPM4,819.9
Rank35GeneCKMEnsemblENSG00000104879nTPM4,663.7
Rank36GenePTGDSEnsemblENSG00000107317nTPM4,605.2
Rank37GeneCOX7A1EnsemblENSG00000161281nTPM4,516.8
Rank38GeneFTH1EnsemblENSG00000167996nTPM4,465.5
Rank39GeneTPT1EnsemblENSG00000133112nTPM4,250.7
Rank40GeneMYL12AEnsemblENSG00000101608nTPM3,965.3
Rank41GeneEEF1A1EnsemblENSG00000156508nTPM3,836.2
Rank42GeneHSPB7EnsemblENSG00000173641nTPM3,567.7
Rank43GeneRPS27EnsemblENSG00000177954nTPM3,392
Rank44GeneHSPB1EnsemblENSG00000106211nTPM3,366.9
Rank45GeneHBA2EnsemblENSG00000188536nTPM3,042.6
Rank46GeneRPL41EnsemblENSG00000229117nTPM2,937.5
Rank47GeneS100A1EnsemblENSG00000160678nTPM2,882.1
Rank48GeneMGPEnsemblENSG00000111341nTPM2,767.3
Rank49GeneRPL10EnsemblENSG00000147403nTPM2,678
Rank50GeneTMSB4XEnsemblENSG00000205542nTPM2,594.9
Rank51GeneALDOAEnsemblENSG00000149925nTPM2,567.3
Rank52GeneRPL37AEnsemblENSG00000197756nTPM2,554.4
Rank53GeneHBBEnsemblENSG00000244734nTPM2,550.9
Rank54GeneFTLEnsemblENSG00000087086nTPM2,480.8
Rank55GeneGSNEnsemblENSG00000148180nTPM2,454.2
Rank56GeneCSRP3EnsemblENSG00000129170nTPM2,447.5
Rank57GeneDCNEnsemblENSG00000011465nTPM2,388
Rank58GenePLA2G2AEnsemblENSG00000188257nTPM2,383.2
Rank59GeneGPX3EnsemblENSG00000211445nTPM2,295.6
Rank60GeneB2MEnsemblENSG00000166710nTPM2,179.2
Rank61GeneMYL9EnsemblENSG00000101335nTPM2,176.3
Rank62GeneATP5MEEnsemblENSG00000169020nTPM2,120.6
Rank63GeneRPS29EnsemblENSG00000213741nTPM2,067.5
Rank64GeneRPLP1EnsemblENSG00000137818nTPM2,024.6
Rank65GeneRPS18EnsemblENSG00000231500nTPM2,023.1
Rank66GeneATP5F1BEnsemblENSG00000110955nTPM2,013.2
Rank67GeneRPL21EnsemblENSG00000122026nTPM1,941.5
Rank68GeneRPS20EnsemblENSG00000008988nTPM1,938.2
Rank69GeneRPL31EnsemblENSG00000071082nTPM1,905.7
Rank70GeneRPS11EnsemblENSG00000142534nTPM1,905.4
Rank71GeneRPL39EnsemblENSG00000198918nTPM1,853.5
Rank72GeneRPS24EnsemblENSG00000138326nTPM1,837.2
Rank73GeneLDHBEnsemblENSG00000111716nTPM1,824.5
Rank74GeneGNASEnsemblENSG00000087460nTPM1,819.8
Rank75GeneRPS12EnsemblENSG00000112306nTPM1,806.2
Rank76GeneBSGEnsemblENSG00000172270nTPM1,787.2
Rank77GeneCOX4I1EnsemblENSG00000131143nTPM1,780.9
Rank78GeneRPS25EnsemblENSG00000118181nTPM1,752.1
Rank79GeneTMSB10EnsemblENSG00000034510nTPM1,700.5
Rank80GeneCOX5AEnsemblENSG00000178741nTPM1,666.8
Rank81GeneRPL26EnsemblENSG00000161970nTPM1,666.4
Rank82GeneNDUFA1EnsemblENSG00000125356nTPM1,658.3
Rank83GeneRPL30EnsemblENSG00000156482nTPM1,642.3
Rank84GeneRPLP2EnsemblENSG00000177600nTPM1,642.2
Rank85GeneIFITM3EnsemblENSG00000142089nTPM1,621.8
Rank86GeneMYBPC3EnsemblENSG00000134571nTPM1,615.9
Rank87GeneRPL13EnsemblENSG00000167526nTPM1,598.5
Rank88GeneCHCHD10EnsemblENSG00000250479nTPM1,596.3
Rank89GeneCOX7CEnsemblENSG00000127184nTPM1,588.4
Rank90GeneUBBEnsemblENSG00000170315nTPM1,562.7
Rank91GeneRPS17EnsemblENSG00000182774nTPM1,562.1
Rank92GeneRPL13AEnsemblENSG00000142541nTPM1,555.1
Rank93GeneRPS16EnsemblENSG00000105193nTPM1,503.1
Rank94GeneLGALS1EnsemblENSG00000100097nTPM1,492.7
Rank95GeneRPS3AEnsemblENSG00000145425nTPM1,482.9
Rank96GeneRPL17EnsemblENSG00000265681nTPM1,474.5
Rank97GeneCOX6B1EnsemblENSG00000126267nTPM1,471.6
Rank98GenePLNEnsemblENSG00000198523nTPM1,463
Rank99GeneRPL19EnsemblENSG00000108298nTPM1,462.9
Rank100GeneRPL35EnsemblENSG00000136942nTPM1,438.2
Rank101GeneATP5F1AEnsemblENSG00000152234nTPM1,408.7
Rank102GeneRPL27AEnsemblENSG00000166441nTPM1,408.1
Rank103GeneRPL38EnsemblENSG00000172809nTPM1,404.5
Rank104GeneFABP3EnsemblENSG00000121769nTPM1,397.8
Rank105GeneNMRK2EnsemblENSG00000077009nTPM1,378.3
Rank106GeneS100A6EnsemblENSG00000197956nTPM1,372.5
Rank107GeneRPL27EnsemblENSG00000131469nTPM1,370.2
Rank108GeneFHL1EnsemblENSG00000022267nTPM1,368.4
Rank109GeneHSPB6EnsemblENSG00000004776nTPM1,366.9
Rank110GeneRPS19EnsemblENSG00000105372nTPM1,357.2
Rank111GeneRPS6EnsemblENSG00000137154nTPM1,354.9
Rank112GeneHBA1EnsemblENSG00000206172nTPM1,348
Rank113GeneFHL2EnsemblENSG00000115641nTPM1,337.6
Rank114GeneRPL3EnsemblENSG00000100316nTPM1,334.5
Rank115GenePSAPEnsemblENSG00000197746nTPM1,333.8
Rank116GeneEEF2EnsemblENSG00000167658nTPM1,329
Rank117GeneACTBEnsemblENSG00000075624nTPM1,326.2
Rank118GeneCST3EnsemblENSG00000101439nTPM1,324.6
Rank119GeneVIMEnsemblENSG00000026025nTPM1,321
Rank120GeneNDUFB1EnsemblENSG00000183648nTPM1,307.9
Rank121GeneRPL24EnsemblENSG00000114391nTPM1,305
Rank122GeneAPODEnsemblENSG00000189058nTPM1,303.8
Rank123GeneRPLP0EnsemblENSG00000089157nTPM1,301.2
Rank124GeneNDUFA13EnsemblENSG00000186010nTPM1,294.7
Rank125GeneNDUFS5EnsemblENSG00000168653nTPM1,286.1
Rank126GeneUBCEnsemblENSG00000150991nTPM1,272.9
Rank127GeneRPL7EnsemblENSG00000147604nTPM1,272.5
Rank128GeneMDH1EnsemblENSG00000014641nTPM1,262.4
Rank129GeneRPL7AEnsemblENSG00000148303nTPM1,260.2
Rank130GeneCOX8AEnsemblENSG00000176340nTPM1,250.2
Rank131GeneDKK3EnsemblENSG00000050165nTPM1,240
Rank132GeneUQCRQEnsemblENSG00000164405nTPM1,218.1
Rank133GeneRPS15AEnsemblENSG00000134419nTPM1,213.4
Rank134GeneRPS15EnsemblENSG00000115268nTPM1,209.7
Rank135GeneRPL18AEnsemblENSG00000105640nTPM1,206.4
Rank136GeneRPL32EnsemblENSG00000144713nTPM1,203.1
Rank137GeneRPL9EnsemblENSG00000163682nTPM1,188.1
Rank138GeneATP5MFEnsemblENSG00000241468nTPM1,187.8
Rank139GeneATP5PFEnsemblENSG00000154723nTPM1,180.7
Rank140GeneRPL23EnsemblENSG00000125691nTPM1,176.2
Rank141GeneRPS8EnsemblENSG00000142937nTPM1,174.2
Rank142GeneRPS2EnsemblENSG00000140988nTPM1,171.4
Rank143GeneRPL23AEnsemblENSG00000198242nTPM1,171
Rank144GeneLDB3EnsemblENSG00000122367nTPM1,170.6
Rank145GeneTAGLNEnsemblENSG00000149591nTPM1,170.5
Rank146GeneATP5MKEnsemblENSG00000173915nTPM1,164.9
Rank147GeneRPS23EnsemblENSG00000186468nTPM1,159.5
Rank148GenePAMEnsemblENSG00000145730nTPM1,159
Rank149GeneSERF2EnsemblENSG00000140264nTPM1,158.8
Rank150GeneATP5POEnsemblENSG00000241837nTPM1,158.4
Rank151GeneCKMT2EnsemblENSG00000131730nTPM1,157.4
Rank152GeneCRIP2EnsemblENSG00000182809nTPM1,152.2
Rank153GeneCD63EnsemblENSG00000135404nTPM1,149.9
Rank154GeneATP5MC3EnsemblENSG00000154518nTPM1,149.4
Rank155GeneCOX5BEnsemblENSG00000135940nTPM1,143.7
Rank156GeneSELENOWEnsemblENSG00000178980nTPM1,129.8
Rank157GeneFAUEnsemblENSG00000149806nTPM1,124.8
Rank158GeneRPS10EnsemblENSG00000124614nTPM1,123.5
Rank159GeneRPL15EnsemblENSG00000174748nTPM1,122.1
Rank160GeneTIMP1EnsemblENSG00000102265nTPM1,120
Rank161GeneRPL28EnsemblENSG00000108107nTPM1,119.1
Rank162GeneRPS13EnsemblENSG00000110700nTPM1,114.3
Rank163GenePGAM2EnsemblENSG00000164708nTPM1,112.4
Rank164GeneRPS9EnsemblENSG00000170889nTPM1,104.4
Rank165GeneUQCRHEnsemblENSG00000173660nTPM1,099.8
Rank166GeneEEF1GEnsemblENSG00000254772nTPM1,097.6
Rank167GeneMYL6EnsemblENSG00000092841nTPM1,096.7
Rank168GeneMT2AEnsemblENSG00000125148nTPM1,093.4
Rank169GeneIGFBP7EnsemblENSG00000163453nTPM1,074.9
Rank170GeneSLC25A3EnsemblENSG00000075415nTPM1,073.6
Rank171GeneRPL34EnsemblENSG00000109475nTPM1,045
Rank172GeneRPL29EnsemblENSG00000162244nTPM1,031.7
Rank173GeneRPL12EnsemblENSG00000197958nTPM1,029.4
Rank174GeneACTA2EnsemblENSG00000107796nTPM1,015.4
Rank175GeneRPS27AEnsemblENSG00000143947nTPM1,013.7
Rank176GeneCOX7BEnsemblENSG00000131174nTPM1,007.2
Rank177GeneCASQ2EnsemblENSG00000118729nTPM989.2
Rank178GeneTPM2EnsemblENSG00000198467nTPM969.6
Rank179GeneRPS14EnsemblENSG00000164587nTPM967.8
Rank180GeneATP5MC2EnsemblENSG00000135390nTPM949.8
Rank181GeneATP2A2EnsemblENSG00000174437nTPM941.9
Rank182GeneUBA52EnsemblENSG00000221983nTPM941.6
Rank183GeneRPL18EnsemblENSG00000063177nTPM938.8
Rank184GeneHSPA1AEnsemblENSG00000204389nTPM935.3
Rank185GeneSLNEnsemblENSG00000170290nTPM931.4
Rank186GeneATP5PDEnsemblENSG00000167863nTPM922.8
Rank187GeneNACAEnsemblENSG00000196531nTPM912
Rank188GeneNDUFB2EnsemblENSG00000090266nTPM907.8
Rank189GeneRPL35AEnsemblENSG00000182899nTPM907.1
Rank190GeneRPL8EnsemblENSG00000161016nTPM905
Rank191GeneCD81EnsemblENSG00000110651nTPM901.7
Rank192GeneRPS7EnsemblENSG00000171863nTPM892.4
Rank193GeneRPS21EnsemblENSG00000171858nTPM885.4
Rank194GeneUQCRBEnsemblENSG00000156467nTPM883.8
Rank195GeneTOMM7EnsemblENSG00000196683nTPM870.1
Rank196GeneRPL5EnsemblENSG00000122406nTPM856.4
Rank197GeneRPL6EnsemblENSG00000089009nTPM855.4
Rank198GeneRPL4EnsemblENSG00000174444nTPM854.7
Rank199GeneCFDEnsemblENSG00000197766nTPM839.2
Rank200GeneMRPL41EnsemblENSG00000182154nTPM826.1
Rank201GenePPIAEnsemblENSG00000196262nTPM810.3
Rank202GeneRPL10AEnsemblENSG00000198755nTPM803.1
Rank203GeneHSPA8EnsemblENSG00000109971nTPM801.5
Rank204GeneBGNEnsemblENSG00000182492nTPM791.1
Rank205GeneNDUFB9EnsemblENSG00000147684nTPM775.4
Rank206GeneRPS5EnsemblENSG00000083845nTPM771.3
Rank207GeneECH1EnsemblENSG00000104823nTPM769.1
Rank208GeneIGKCEnsemblENSG00000211592nTPM761.6
Rank209GeneRACK1EnsemblENSG00000204628nTPM758.2
Rank210GeneNDUFB10EnsemblENSG00000140990nTPM755.8
Rank211GeneHADHBEnsemblENSG00000138029nTPM749.3
Rank212GeneFBLN1EnsemblENSG00000077942nTPM748.5
Rank213GeneHAMPEnsemblENSG00000105697nTPM748.2
Rank214GeneRPSAEnsemblENSG00000168028nTPM746.7
Rank215GeneHSP90AB1EnsemblENSG00000096384nTPM734.1
Rank216GeneLMOD2EnsemblENSG00000170807nTPM728.6
Rank217GeneBMP10EnsemblENSG00000163217nTPM727.2
Rank218GeneCOX6CEnsemblENSG00000164919nTPM726.8
Rank219GeneEEF1A2EnsemblENSG00000101210nTPM722.2
Rank220GeneMYOZ2EnsemblENSG00000172399nTPM722
Rank221GeneMGST3EnsemblENSG00000143198nTPM721.3
Rank222GeneTXNIPEnsemblENSG00000265972nTPM710
Rank223GeneSMPXEnsemblENSG00000091482nTPM709.4
Rank224GeneCOX7A2EnsemblENSG00000112695nTPM706
Rank225GenePRDX2EnsemblENSG00000167815nTPM704.2
Rank226GeneYBX1EnsemblENSG00000065978nTPM703.4
Rank227GeneATP5MC1EnsemblENSG00000159199nTPM699.4
Rank228GeneACTN2EnsemblENSG00000077522nTPM695.9
Rank229GeneMPC1EnsemblENSG00000060762nTPM692.4
Rank230GeneHRCEnsemblENSG00000130528nTPM692.1
Rank231GeneRPS3EnsemblENSG00000149273nTPM691.4
Rank232GeneTPI1EnsemblENSG00000111669nTPM688.7
Rank233GenePDE4DIPEnsemblENSG00000178104nTPM686.9
Rank234GeneGABARAPEnsemblENSG00000170296nTPM686.8
Rank235GeneRPS26EnsemblENSG00000197728nTPM683.8
Rank236GeneRPL36EnsemblENSG00000130255nTPM681.8
Rank237GeneUBL5EnsemblENSG00000198258nTPM674.4
Rank238GeneATP5F1CEnsemblENSG00000165629nTPM671.9
Rank239GeneRPL11EnsemblENSG00000142676nTPM668.8
Rank240GeneHSPA1BEnsemblENSG00000204388nTPM667.7
Rank241GeneNR4A1EnsemblENSG00000123358nTPM661.1
Rank242GeneACTG1EnsemblENSG00000184009nTPM659.8
Rank243GeneNDUFB7EnsemblENSG00000099795nTPM659.6
Rank244GeneCAPNS1EnsemblENSG00000126247nTPM657.9
Rank245GeneCD59EnsemblENSG00000085063nTPM656.7
Rank246GeneEIF3KEnsemblENSG00000178982nTPM638.5
Rank247GenePKMEnsemblENSG00000067225nTPM633.2
Rank248GeneNDUFA11EnsemblENSG00000174886nTPM632
Rank249GeneIFITM2EnsemblENSG00000185201nTPM628.5
Rank250GeneSKP1EnsemblENSG00000113558nTPM614.4

The rank is the row's place in this cut of the table, largest value first, and is not a figure the atlas publishes. The values are the atlas's, sorted by the atlas's own numbers.

  • Human Protein Atlas, the consensus tissue table, the rows for heart muscle · Human Protein Atlas version 25.1 (release 2026-05-25); entry tag 25, rna_tissue_consensus.tsv 2025-11-06 · read · the atlas's consensus tableHuman Protein Atlas version 25.1 (release 2026-05-25); entry tag 25, proteinatlas.org; data available from v25.proteinatlas.org/download/tsv/rna_tissue_consensus.tsv.zip (Uhlén M et al. Science 2015). CC BY 4.0.